Is it possible to "un-pause gameplay" during an action list and have it continue? If I have "When Running: Pause Gameplay" set for an action list, is it possible to set gameplay to "Run in Background" from the actionlist for the remainder of the actions in the list?
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No - you'll need to split it into two separate ActionLists.
Just to re-confirm, "Pause Gameplay" mode is basically just hiding the cursor, or does it change any other systems?
No, it disables a lot - movement, interactions, certain inputs, menus and more.
You can, however, use the Engine: Manage systems Action to selectively disable certain systems during gameplay - so you could possibly have the ActionList run in the background, and disable gameplay systems for part of it.
This seems to do the trick, is there a way I could make a custom action version of this that only has the two parts needed, since it is enormous in the graph for just hiding the cursor and disabling input:
https://imgur.com/WjnmaqY
You can collapse the Action to reduce its size, but you could create a custom Action by duplicating the original (ActionSystemLock.cs) and removing the undesired fields/variables.
The Cursor and Input fields rely on the cursorLock and inputLock variables - all else can be removed.